Species Habitat Index

Overview

The integrity of ecosystems is broadly defined by the status of their component species and the ecological processes they support and require. Integrity can be assessed by the degree of change (loss and gain) in the set of species and associated processes observed within an ecosystem and its habitats. The Species Habitat Index (SHI) measures this change and captures alterations in the ecological intactness of ecosystems. The SHI is calculated annually across tens of thousands of terrestrial vertebrate species.

The index measures changes in the estimated size, connectivity and quality of species habitats. The index uses species as core units of analysis, thereby capturing the individual ecological processes associated with species that are central to ecosystem integrity. The index uses spatially explicit information at a resolution of single pixels (1 km2) to support aggregate measures of the ecological integrity of defined geographic units. Given the species-level nature of the metric, SHI informs about trends in species population size, distribution, health, and, as proxy, genetic diversity.

The SHI was developed under the auspices of the GEO Biodiversity Observation Network and part of the Biodiversity Indicators Partnership and is formally adopted in the Global Biodiversity Monitoring Framework as a component indicator for Goal A, which stipulates that the "integrity, connectivity and resilience of all ecosystems" be "maintained, enhanced, or restored" by 2050.

The integrity of ecosystems is broadly defined by the status of their component species and the ecological processes they support and require. Integrity can be assessed by the degree of change (loss and gain) in the set of species and associated processes observed within an ecosystem and its habitats. The Species Habitat Index (SHI) measures this change and captures alterations in the ecological intactness of ecosystems. The SHI is calculated annually across tens of thousands of terrestrial vertebrate species.

The index measures changes in the estimated size, connectivity and quality of species habitats. The index uses species as core units of analysis, thereby capturing the individual ecological processes associated with species that are central to ecosystem integrity. The index uses spatially explicit information at a resolution of single pixels (1 km2) to support aggregate measures of the ecological integrity of defined geographic units. Given the species-level nature of the metric, SHI informs about trends in species population size, distribution, health, and, as proxy, genetic diversity.

A METRIC OF

ECOSYSTEM INTEGRITY

COMPONENT INDCATOR OF

GOAL A

TAXONOMIC COVERAGE

TERRESTRIAL:

The SHI was developed under the auspices of the GEO Biodiversity Observation Network and part of the Biodiversity Indicators Partnership and is formally adopted in the Global Biodiversity Monitoring Framework as a component indicator for Goal A, which stipulates that the "integrity, connectivity and resilience of all ecosystems" be "maintained, enhanced, or restored" by 2050.

Media & Presentations

Methods

Habitat-suitable range

We apply statistical models incorporating species expert ranges, habitat preferences, environmental layers, and species occurrence points to develop species habitat-suitable range maps. These maps represent the habitats within a species range boundaries where the species is most likely to inhabit and therefore provide a more accurate representation of the size and distribution of the species range.

Species-level scores

The SHI calculation is based on individual species habitat scores (SHS). The SHS measures change over time in the area and connectivity of species’ habitat-suitable ranges using 2001 as the baseline year. The SHS is given by the average of the area score and the connectivity score.

The area score measures changes in the size of the species’ habitat-suitable range. This is given by the product of summed suitability (continuous range of 0 – 1) of individual landscape pixels and their size. For suitability expressed in binary form (presence-absence maps) for 1 km2 pixel, this is simply the total presence pixel count.

The connectivity score measures changes in the average connectedness, or fragmentation, of the species’ habitat-suitable range. This is given as the average distance to the edge of the suitable area across all suitable pixels (a widely-used, robust measure of connectivity - GISfrag metric). For custom calculations at national level this can be extended to include other information, e.g. measures that weight the distance among habitats by the resistance to movement of the intervening landscape.

The area and connectivity scores are given as the change since 2001, where the baseline is represented as SHS = 100. Thus, a loss of 6% of habitat area would give an area score of 94 and a loss of 4% of connectivity would give a connectivity score of 96. The SHS is the simple average of these two scores, so this species would have an SHS of 95 compared to the reference year 2001.

National SHI

The SHI of a country or other geographic unit is given as the average SHS of all species within that unit. SHI values for a country can either be computed as the simple mean across species (National SHI) or by weighting species-level values by the proportion of the global population the country is estimated to hold (Steward’s SHI). In addition to reporting on the separate Area or Connectivity aspects of SHI, indicator subsets can address different species groups, e.g. species dependent on certain habitats and ecosystems, rare or threatened species, or those with particularly rapid recent habitat changes.

Sources

For the latest 2023 version of the SHI:

Species Range Maps

Species Group

Source

Amphibians

IUCN (2016). The IUCN Red List of Threatened Species. International Union for Conservation of Nature. Accessed on January 2017. Downloaded at www.iucnredlist.org

Birds

Jetz, W., et al. (2012). The global diversity of birds in space and time. Nature, (491);444-448. doi.org/10.1038/nature11631

Mammals

Mammal Diversity Database. (2020). Mammal Diversity Database (Version 1.2) [Data set]. Zenodo. doi.org/10.5281/zenodo.4139818. Map of Life. (2021). Mammal range maps harmonised to the Mammals Diversity Database [Data set]. Map of Life. doi.org/10.48600/MOL-48VZ-P413

Reptiles

Roll, U. and Meiri, S. (2022). GARD 1.7 - updated global distributions for all terrestrial reptiles [Dataset]. Dryad. doi.org/10.5061/dryad.9cnp5hqmb

Terrestrial vertebrates

Misc. literature and expert sources

Species Occurrence Points

All point datasets include data from 2001-02-01 to 2023-12-31.

Species Group

Source

Terrestrial vertebrates

GBIF.org (1 June 2022) GBIF Occurrence Download: https://doi.org/10.15468/dl.4hpkfz

Birds

eBird Basic Dataset. Version: EBD_relMay-2022. Cornell Lab of Ornithology, Ithaca, New York. May 2022.

Species Habitat Preferences

Species Group

Source

Terrestrial vertebrates

IUCN Red List of Threatened Species. 2025. Version 2025-2. www.iucnredlist.org. Downloaded on 2025-04-22.

Terrestrial vertebrates

Misc. literature and expert sources

Environmental Layers

Dataset

Source

Elevation

Amatulli, G., et al. (2018) A suite of global, cross-scale topographic variables for environmental and biodiversity modeling. Scientific Data (5);180040. https://doi.org/10.1038/sdata.2018.40. Available at www.earthenv.org/topography.

Land cover

ESA. Land Cover CCI Product User Guide Version 2. Tech. Rep. (2017). Version 2022. Available at: maps.elie.ucl.ac.be/CCI/viewer/download/ESACCI-LC-Ph2-PUGv2_2.0.pdf

Tree cover

Hansen, M. C., et al. (2013) High-Resolution Global Maps of 21st-Century Forest Cover Change. Science 342 (15 November): 850-53. Version 2023. Data available online from: glad.earthengine.app/view/global-forest-change.

Region Layers

Dataset

Source

Country boundaries

Database of Global Administrative Boundaries (GADM) version 4.1. Available online at gadm.org/data.htm.

Citations & Acknowledgements

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Supported by

Winner of

Stay in the know.

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600 N Broad Street Suite 5 #4038 Middletown, DE 19709

Copyright © 2025

Map of Life

Research by

Supported by

Winner of

Stay in the know.

Connect with us

600 N Broad Street Suite 5 #4038 Middletown, DE 19709

Copyright © 2025